1College of Horticulture, Nanjing Agricultural University, Nanjing, Jiangsu, 210095, China.
2Institute of Botany, Jiangsu Province and the Chinese Academy of Sciences, Nanjing 210014, China.
3Key Laboratory for Forest Resources Conservation and Use in the Southwest Mountains of China, Ministry of Education, Southwest Forestry University, Kunming, 650224, China.
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